best mid-layer score is +0.052285 +/- 0.000040 and is always achieved at species_count 144, coherence_noise 0.7, pulse_layer 1; for species_count 96 and coherence_noise 0.7, pulsing the middle layer raises area gain by +0.06209 +/- 0.00037 and retention by +0.14809 +/- 0.00008 relative to basal, whereas pulsing the top layer at the same noise drives area gain negative to -0.33199 +/- 0.00031.
The food-web response is genuinely layer selective. Mid-layer pulsing behaves like a band-pass filter: it preserves or enlarges response area while also improving retention, whereas top-layer forcing over-attenuates the signal and collapses area. The strong noise dependence suggests that intermediate coherence is needed for the middle trophic layer to redistribute pulses without simply dissipating them. The implication is that trophic position, not just pulse magnitude, determines whether perturbations are filtered, amplified, or lost as they propagate through the stack.
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